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Harley Quinn joins forces with Poison Ivy and Catwoman in the series Gotham City Sirens. Having moved in with Pamela Isley at the Riddler’s apartment, she meets up with Catwoman, who offers for the three of them to live and work together. A new villain who tried to take down Selina Kyle named Boneblaster breaks into the apartment, and the three of them have to move after they defeat him. Later, after a chance encounter with Hush, the Joker attempts to kill her, apparently out of jealousy. Quinn is rescued by Ivy and Catwoman, and it is later revealed that her attacker wasn’t the real Joker, but one of his old henchmen impersonating him.

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In Birds of Prey #105, Harley Quinn is revealed as the sixth member of the Secret Six. In issue #108, upon hearing that Oracle has sent the Russian authorities footage of teammate Deadshot murdering the Six’s employer as payback for double-crossing them, Harley asks, “Is it a bad time to say ‘I quit’?”, thus leaving the team.
In Countdown #43, Harley appears to have reformed and is shown to be residing in an Amazon-run women’s shelter. Having abandoned her jester costume and clown make-up, she now only wears an Amazonian stola or chiton. She befriends the former Catwoman replacement Holly Robinson, and then succeeds in persuading her to join her at the shelter, where she is working as an assistant. They are both brought to Themiscyra by “Athena” (really Granny Goodness) and begin Amazon training. Holly and Harley then meet the real Athena, and encounter Mary Marvel. The group reveal Granny’s deception, and Holly, Harley, and Mary follow her as she retreats to Apokolips. Mary finds the Olympian gods, whom Granny had been holding prisoner, and the group frees them. Harley is granted powers by Thalia as a reward. Upon returning to Earth, the powers vanish, and Harley and Holly return to Gotham City.

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In the One Year Later continuity, Harley Quinn is an inmate at Arkham, glimpsed briefly in Detective Comics #823.
Harley next appeared in Batman #663, in which she helps the Joker with a plan to kill all his former henchmen, unaware that the “punch line” to the scheme is her own death. Upon realizing this, she shoots him in the shoulder.
Harley resurfaces in Detective Comics #831, written by Paul Dini. Harley has spent the last year applying for parole, only to see her request systematically rejected by Bruce Wayne, the layman member of Arkham’s medical commission. She is kidnapped by Peyton Riley, the new female Ventriloquist, who offers her a job; Harley turns the job down out of respect for the memory of Arnold Wesker, the original Ventriloquist, who attempted to cheer her up during her first week in Arkham while the Joker was still on the loose. She then helps Batman and Commissioner Gordon foil the impostor’s plans. Although Riley escapes, Bruce Wayne is impressed with Harley’s effort at redemption, and agrees with granting her parole.

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Quinn’s DC Universe comic book origin, revealed in Batman: Harley Quinn (October 1999), is largely an adaptation of her animated origin from the Mad Love graphic novel.
A Harley Quinn ongoing series was published monthly by DC Comics for 38 issues from 2001 to 2003. Creators who contributed to the title included Karl Kesel, Terry Dodson, A.J. Lieberman, and Mike Huddleston. The series dealt with her going solo, eventually starting a gang and then fleeing Gotham for the city of Metropolis with her friend Poison Ivy. Quinn dies, only to be resurrected and return to Gotham. The series ends with Harley turning herself in to Arkham Asylum, having finally understood that she needs help. We also learn in issue #8 of the comic that Harley had a relationship in college with fellow psychiatry major Guy Kopski whose suicide started her obsession with the Joker. Harley later appears in the Jeph Loeb series Hush. She is next seen in a Villains United Infinite Crisis special, where she is one of the many villains who escape from Arkham (although she is knocked unconscious the moment she escapes).

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The character proved so popular that she was eventually added to the Batman comic book canon (although she had already appeared in the Elseworlds Batman: Thrillkiller and Batman: Thrillkiller ’62 in 1997). The comic book version of Quinn, like the comic book version of the Joker, is more dangerously violent and less humorously quirky than the animated series version. Despite her noticeably more violent demeanor, Harley does show mercy and compassion from time to time; she notably stops Poison Ivy from killing Batman, instead convincing her to leave the hero hanging bound and gagged from a large statue. Batman is later untied by Batgirl.

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She appeared in World’s Finest: The Batman/Superman Movie as a rival and foil for Lex Luthor’s assistant Mercy Graves; each takes an immediate dislike for the other, at one point fighting brutally with each other as Lex Luthor and the Joker have a business meeting. In the film’s climax, Harley nearly kills Graves when she gags her and ties her to pole on an island that is going to explode within an hour, but Superman finds her and unties her.
The animated movie Batman Beyond: Return of the Joker takes place in the future, long after the events in Batman: The Animated Series. It includes a flashback scene with Harley falling down a deep pit during a battle with Batgirl. At the end of the movie, a pair of twin juvenile girls who model themselves on the Joker are released on bail to their grandmother, who angrily berates them—to which they answer: “Oh, shut up, Nana Harley!”

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After Batman: The Animated Series and The New Batman Adventures, Harley makes several other animated appearances. She appears as one of the four main female characters of the web cartoon Gotham Girls. She also made guest appearances in other cartoons in the DC animated universe, appearing in the Justice League episode “Wild Cards” (alongside the Joker) and the Static Shock episode “Hard as Nails” (alongside Poison Ivy).